They are an alternative to the tailscale operated DERP servers, which are cloud relays.
Even with the much touted NAT punching capabilities of tailscale, there are numerous instances where tailscale cannot establish a true p2p connection. The last fallback is the quite slow DERP relay and from experience it gets used very often.
If you have a peer in your tailscale network that has a good connection and that maybe you can even expose to the internet with a port forward on your router, you now have this relay setting that you can enable to avoid using the congested/shared DERP servers. So there is not really a new use-case for this. It's the same, just faster.
Even with the much touted NAT punching capabilities of tailscale, there are numerous instances where tailscale cannot establish a true p2p connection. The last fallback is the quite slow DERP relay and from experience it gets used very often.
If you have a peer in your tailscale network that has a good connection and that maybe you can even expose to the internet with a port forward on your router, you now have this relay setting that you can enable to avoid using the congested/shared DERP servers. So there is not really a new use-case for this. It's the same, just faster.