Is this novel? Spatial codes have been known for long and neuron (populations) are known to code for time [1]. That reference also states that "Neurons that respond selectively to the temporal and spatio-temporal structure of sensory stimuli have also been identified in the auditory system of birds and mammals". Doesn't that pre-empt the conclusion that "Thus, the brain can represent time and space as overlapping but dissociable dimensions".
[1]: https://royalsocietypublishing.org/doi/10.1098/rstb.2012.046...