This is fascinating. Naughty Dog has developed for Sony consoles since Crash, so they might have low visibility if you're not in touch with the Playstation ecosystem, but they're renowned for their total mastery of successive generations of the Playstation hardware.
HNers might be sad to learn that they gave up writing games in their home-grown Lisp with the advent of the PS3 era. It was called GOOL, and later, GOAL: http://en.wikipedia.org/wiki/Game_Oriented_Assembly_Lisp