The whole point of the article is that the traditional "calories in/calories out" model is wrong.
"Based on the research, Pontzer has proposed a new model that upends the the old "calories in, calories out" approach to exercise... He calls this the "constrained model" of energy expenditure, which shows that the effect of more physical activity on the human body is not linear."
"Based on the research, Pontzer has proposed a new model that upends the the old "calories in, calories out" approach to exercise... He calls this the "constrained model" of energy expenditure, which shows that the effect of more physical activity on the human body is not linear."